The Lion (Panthera leo) is one of four big cats in the genus Panthera, and a member of the family Felidae. With some males exceeding 250 kg (550 lb) in weight,[4] it is the second-largest living cat after the tiger. Wild lions currently exist in Sub-Saharan Africa and in Asia with a critically endangered remnant population in northwest India, having disappeared from North Africa, the Middle East, and Western Asia in historic times. Until the late Pleistocene, which was about 10,000 years ago, the lion was the most widespread large land mammal after humans. They were found in most of Africa, much of Eurasia from western Europe to India, and in the Americas from the Yukon to Peru.
Lions live for around 10–14 years in the wild, while in captivity they can live over 20 years. In the wild, males seldom live longer than ten years as fights with rivals occasionally cause injuries.[5] They typically inhabit savanna and grassland, although they may take to bush and forest. Lions are unusually social compared to other cats. A pride of lions consists of related females and offspring and a small number of adult males. Groups of female lions typically hunt together, preying mostly on large ungulates. The lion is an apex and keystone predator, although they will scavenge if the opportunity arises. While lions do not typically hunt humans selectively, some have been known to become man-eaters and seek human prey.
The Lion is a vulnerable species, having seen a possibly irreversible population decline of 30 to 50 percent over the past two decades in its African range.[6] Lion populations are untenable outside of designated reserves and national parks. Although the cause of the decline is not fully understood, habitat loss and conflicts with humans are currently the greatest causes of concern. Lions have been kept in menageries since Roman times and have been a key species sought for exhibition in zoos the world over since the late eighteenth century. Zoos are cooperating worldwide in breeding programs for the endangered Asiatic subspecies.
Visually, the male lion is highly distinctive and is easily recognized by its mane. The lion, particularly the face of the male, is one of the most widely recognized animal symbols in human culture. Depictions have existed from the Upper Paleolithic period, with carvings and paintings from the Lascaux and Chauvet Caves, through virtually all ancient and medieval cultures where they historically occurred. It has been extensively depicted in literature, in sculptures, in paintings, on national flags, and in contemporary films and literature.
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Etymology
The lion's name, similar in many Romance languages, derives from the Latin leo;[7] cf. the Ancient Greek λέων (leon).[8] The Hebrew word לָבִיא (lavi) may also be related,[9] as well as the Ancient Egyptian rw.[10] It was one of the many species originally described, as Felis leo, by Linnaeus in his eighteenth century work, Systema Naturae.[3] The generic component of its scientific designation, Panthera leo, often is presumed to derive from Greek pan- ("all") and ther ("beast"), but this may be a folk etymology. Although it came into English through the classical languages, panthera is probably of East Asian origin, meaning "the yellowish animal," or "whitish-yellow".[11]
Taxonomy and evolution
The oldest lion-like fossil is known from Laetoli in Tanzania and is perhaps 3.5 million years old; some scientists have identified the material as Panthera leo. These records are not well-substantiated, and all that can be said is that they pertain to a Panthera-like felid. The oldest confirmed records of Panthera leo in Africa are about 2 million years younger.[12] The closest relatives of the lion are the other Panthera species: the tiger, the jaguar, and the leopard. Morphological and genetic studies reveal that the tiger was the first of these recent species to diverge. About 1.9 million years ago the jaguar branched off the remaining group, which contained ancestors of the leopard and lion. The lion and leopard subsequently separated about 1 to 1.25 million years ago from each other.[13]
Panthera leo itself evolved in Africa between 1 million and 800,000 years ago, before spreading throughout the Holarctic region.[14] It appeared in Europe for the first time 700,000 years ago with the subspecies Panthera leo fossilis at Isernia in Italy. From this lion derived the later Cave Lion (Panthera leo spelaea), which appeared about 300,000 years ago. During the upper Pleistocene the lion spread to North and South America, and developed into Panthera leo atrox, the American Lion.[15] Lions died out in northern Eurasia and America at the end of the last glaciation, about 10,000 years ago;[16] this may have been secondary to the extinction of Pleistocene megafauna.[17]
Subspecies
Traditionally, twelve recent subspecies of lion were recognized, the largest of which has been recognized as the Barbary Lion.[18] The major differences separating these subspecies are location, mane appearance, size, and distribution. Because these characteristics are very insignificant and show a high individual variability, most of these forms were debatable and probably invalid; additionally, they often were based upon zoo material of unknown origin that may have had "striking, but abnormal" morphological characteristics.[19] Today only eight subspecies usually are accepted,[16][20] but one of these (the Cape Lion formerly described as Panthera leo melanochaita) probably is invalid.[20] Even the remaining seven subspecies might be too many; mitochondrial variation in recent African lions is modest, which suggests that all sub-Saharan lions could be considered a single subspecies, possibly divided in two main clades: one to the west of the Great Rift Valley and the other to the east. Lions from Tsavo in Eastern Kenya are much closer genetically to lions in Transvaal (South Africa), than to those in the Aberdare Range in Western Kenya.[21][22]
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